CamTarn is entirely correct, they are for ventilation.
House building is a constant battle between the needs of the
inhabitants (for
warmth and
comfort), and the needs of the building (
ventilation etc).
If
timbers are not
ventilated, they will quickly
rot. If rooms are not ventilated,
the moisture you give off will not escape, and the
walls will sweat, and the wallpaper will go
mouldy and fall off the walls.
If you have a
gas fire in your room, and it can't get enough oxygen, it will produce
carbon monoxide as a by-product, which will prevent you from enjoying it's warmth for very long.
Overall, you need ventilation all over your house (grates between rooms), and
airbricks to the outside world. However, they should be placed so that
draughts do not cause a problem. If you're ventilating a fire, have the vent on the wall or in the floor near the fire, so that the draught has a minimum distance to travel.
Radiators are ideally placed in places where the vents are, since the
cool air will be warmed immediately, rather than travelling across the room before heing heated.
The advantage of vents is that you can place them where it's
convenient for the required ventilation, and you can then block up all unwanted draughts (for example, those from
under doors and around
windows).