The little dimple that can be found right under your nose. A great place to rest the end of your pointer finger (or a Faber #2 pencil) when thinking "hmm...".

I'm not sure of the exact "reason" for the philtrum, so I guess it's right up there with the appendix — although I've never heard of anyone having to have an emergency filtrumdectomy. Perhaps it provides a clearer path for for air flow during sniffing. I have some time booked at the local wind tunnel to test this theory — wish me luck.

I once mistakenly believed that the word was spelled "filtrum" — luckily geeklizzard enlightened me.